Genus: Cell body short, radiate in shape; apical view of the cell, triangular, polygonal, 
ellipsoidal or spindle-shaped; zygospores formed outside of mother cells 
 
Subgenus: lacks arm-like projections 
(Illustrations of The Japanese Fresh-water Algae, 1977) 
 
Species: 
semicells twisted with each other at isthmus 
(Illustrations of The Japanese Fresh-water Algae, 1977) 
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