Genus: Colonies of a fixed number of cells, flat, circular in shape; cell body polygonal in shape, with horn-like projections; cell wall garanulated, wrinkled or notched; chloroplasts plate-like or reticular; asexual reproduction by zoospore; sexual reproduction by isogametes (Illustrations of The Japanese Fresh-water Algae, 1977).
Species: Colonies of 8, 16, 32 (sometimes 64, 128) cells, spherical or ellipsoid in shape, intercellular space absent; inner cells tetragonal to hexagonal, concaved at outer side; marginal cells with two long horn-like processes; a tuft of bristles projected from apex of the processes; zoospores produced within the mother cells; daughter colonies formed in a vesicle released from the mother cells; cell wall granulated (Photomicrographs of the fresh-water algae, 1993).

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Horn-like processes long
var. boryanum 32, 16 cells 16 = 10+5+1 col. μm φ Japan, 1998	var. boryanum 16 = 10+5+1 col. μm φ Abiko Chiba, 2000	var. boryanum stock Ton-2 16 = 10+5+1 col. μm φ Nishinasuno Tochigi, 2000	var. boryanum stock Ton-2 16 = 10+5+1 col. μm φ Nishinasuno Tochigi, 2000
var. boryanum 16 = 10+5+1 col. μm φ Koshigaya Saitama, 2001	var. boryanum 16 = 7+1 Kenmin-kenko f.m. Koshigaya Saitama, 2006
var. boryanum 32, 16 cells col. μm φ Japan, 199?	var. boryanum 16 cells = 11+5 col. μm φ Shinobazu-ike Tokyo, 2001	var. boryanum 16 cells = 11+5 col. μm φ Saitama, 1999
var. boryanum ? granules large col. μm φ USA, 1999	var. boryanum surface granulated col. μm φ Umi-no-nakamichi Fukuoka, 2001	var. boryanum ? or brevicorne ? stock Tom-10 surface granulated col. μm φ Mizumoto Park Tokyo, 2001	var. boryanum surface granulated col. μm φ Heisei-no-mori Park Kawashima or Kawajima Saitama, 2002
var. boryanum granules large col. μm φ Chiba-Minato Park Chiba, 2003	var. boryanum ? col. μm φ Chiba-Minato Park Chiba, 2003	var. boryanum ? col. μm φ Chiba-Minato Park Chiba, 2003	var. boryanum ? col. μm φ Higusa-numa Hikari Chiba, 2003
var. boryanum ? cell μm Kurome river Asaka Saitama, 2000	var. boryanum col. 35 μm φ Nasuno-ga-hara m. Nishi-nasuno Tochigi, 2004	var. boryanum col. μm φ Kawanago Ishibashi Tochigi, 2004	var. boryanum col. μm φ Iruma river Kawagoe Saitama, 2004
var. boryanum col. 50 μm φ Aquatic botanical g. Sawara Chiba, 2004	var. boryanum col. 57-65 μm φ Takao Sakura p. Kitamoto Saitama, 2005	var. boryanum col. 60 μm φ Shinrin-koen Namegawa Saitama, 2005	var. boryanum col. 50,80 μm φ Lake Shirakaba Tateshina & Chino Nagano, 2005
var. boryanum col. μm φ Obihiro-Hasshonochi Obihiro Hokkaido, 2005	var. boryanum col. μm φ riverside park Abukuma river Fukushima, 2005	var. boryanum col. μm φ Shiroyama park Okegawa Saitama, 2006	var. boryanum col. 75 μm φ Nakayado h.p. Fukaya Saitama, 2006
Horn-like processes short
P. boryanum ? horns very short 16 = 10+5+1 col. μm φ Uruguay, 1999	P. boryanum ? horns very short 32 = 16+10+5+1 col. μm φ Uruguay, 1999	P. boryanum ? stock 37-2D horns very short 16 = 10+5+1 col. μm φ Uruguay, 1999	P. boryanum ? stock 37-2D horns very short 16 = 10+5+1 col. μm φ Uruguay, 1999
P. boryanum ? or P. integrum ? stock 37-2E horns very short 16 = 10+5+1 col. μm φ Uruguay, 1999	P. boryanum ? or P. integrum ? stock 37-2E horns very short 16 = 10+5+1 col. μm φ Uruguay, 1999	P. boryanum ? horns very short Omine-numa Tsukiyono Gunma, 2003	var. boryanum col. 105, 125 μm φ Kumakubo Nasu-shiobara Tochigi, 2005

Similar species -->> P. biradiatum; P. muticum;

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