Genus: Cell body flat, large, variable in shape, longer than wide; strong constriction at center; each semicell with several notches; cell wall with many short spines (Illustrations of The Japanese Fresh-water Algae, 1977).
Species [ var. thomasiana ]: Cell body 95-98 μm long, 97-98 μm wide, isthmus 22-23 μm wide,

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Conspicuous protrusions present at the center of the cell
v. thomasiana 223 x 190 μm Sukawa highland Kurikoma mountain Akita, 2005	v. javanica μm Miyatoko m.4 Minami-Aizu Fukushima, 2008	v. javanica μm Miyatoko m.6 Minami-Aizu Fukushima, 2008	v. javanica μm Kareki-numa Nikko Tochigi, 2008.8
v. javanica μm Kareki-numa Nikko Tochigi, 2008.9	v. pulcherrima 220 x 185 μm Kakuman-buchi Akagi mountain Gunma, 2005
Conspicuous protrusions absent at the center of the cell; similar to M. denticulata v. javanica
v. notata 225 x 205 μm Tashirodaira Hakkoda mountain Aomori, 2006	v. notata 225 x 180 μm Igashira park Mooka Tochigi, 2006	v. notata 284 x 262 μm Ichinose-enchi Norikura highland Nagano, 2006	v. notata μm Dojo-ike Norikura highland Nagano, 2007
v. notata 240 x 205 μm Kowashimizu Suwa Nagano, 2007	v. notata μm Yamakado marsh Nishi-Asai Shiga, 2008	v. notata μm Kashibaru m.2-2 Karatsu Saga, 2008	v. notata μm Kashibaru m.II Karatsu Saga, 2008

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